Fig. 1

Gross images of the grouper fish: A rostral view of the opened mouth and B lateral view of the gills with reflected operculum that explained the structures and position of the gills. (A1, A2, A3, and A4) 1st–4th gill arches. rackers (R), a lateral row of gill rakers (LR), esophagus (O), interbranchial septum (IS), the mandible (M), the roof of the pharynx (RP), the roof of the oral cavity (RO), pharyngeal teeth (PT), chewing pads (Cp), operculum (OP), rostral extremities (RE), caudal extremities (CE), short epibranchial (eb), long ceratobranchial (cb), angle of epi-ceratobranchial union (ag), gill raker (R), gill arch (A), gill filaments (F), and tongue (T)